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Plot: Fisherman Matthew Boyd (Richard Dreyfuss) is fishing in Lake Victoria, AZ when a small earthquake hits, splitting the lake floor and causing a whirlpool. Boyd falls in and is ripped apart by a school of prehistoric piranhas that emerge from the chasm and ascend the vortex.

Jake (McQueen) is admiring attractive tourists as Spring Break begins. He reunites with his old crush, Kelly (Jessica Szohr) and meets Derrick Jones (Jerry O'Connell) an eccentric pornographer, as well as Danni (Kelly Brook), one of his actresses. Derrick convinces Jake to show him good spots on the beach for filming a pornographic movie.

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That night, Jake's mother, Sheriff Julie Forester (Elisabeth Shue), searches for the missing Matthew Boyd with Deputy Fallon (Ving Rhames), who has found Boyd's boat. They find his mutilated body and contemplate closing the beach down. The next morning, a lone cliff diver is attacked and consumed by the marauding fish.

Jake bribes his younger sister and brother, Laura (Brooklynn Proulx) and Zane (Sage Ryan), to stay home alone so that he can show Derrick around the beach with Danni. After Jake leaves, Zane drafts Laura to go fishing on a small sandbar island. They forget to tie the boat down and are stranded in the middle of the lake. Meanwhile, Jake goes to meet with Derrick and runs into Kelly, who invites herself onto Derrick's boat, The Barracuda. Jake meets Crystal (Riley Steele), another of Derrick's actresses, and cameraman Drew (Paul Scheer). They film an underwater love scene between Danni and Crystal before partying.

Meanwhile, Julie takes a team of seismologist divers — Novak (Adam Scott), Sam (Ricardo Chavira), and Paula (Dina Meyer) — to the fissure. Novak speculates that the rift leads to a buried prehistoric lake. Paula and Sam scuba dive to the bottom and discover a large cavern, filled with large egg stalks. Both are killed before they can escape or alert the others. Novak and Julie pull Paula's corpse into the boat, capturing a lone piranha, which they take to Mr. Goodman (Christopher Lloyd), a marine biologist. He explains that the piranhas are a prehistoric species, long believed-extinct, and must had been trapped in the underground lake for over two million years.

Julie, Novak, and Fallon try to evacuate the beach, but their warnings are ignored until the piranhas attack the tourists. Novak boards a jet-ski with a shotgun to help while Fallon ushers people to shore and Julie tries to get swimmers into the police boat. A floating stage set up in the water collapses from the weight of all the panicking guests and the wet T-shirt contest host (Eli Roth) is killed.

Jake spots Laura and Zane on the island, and forces Derrick to rescue them. Derrick gets the boat's propeller stuck and crashes into some rocks, breaking the glass observation dome and flooding the rooms below deck. Kelly is trapped in in the kitchen while Derrick, Crystal, and Drew are thrown from the boat. Crystal and Drew are devoured while Danni manages to get a partially eaten Derrick back on board. Jake, panicking that he can't reach Kelly, calls Julie.

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Production: Chuck Russell was originally scheduled to direct the film, and made uncredited rewrites to the script by Josh Stolberg and Peter Goldfinger, as well as incorporating the original John Sayles script that Joe Dante directed the first time around. Alexandre Aja was selected to direct the film instead. Production on the film was scheduled to begin late 2008, but was delayed until March 2009. In October 2008, director Alexandre Aja stated filming would begin in the spring. He further stated "it's such a difficult movie, not only because of the technicality of it and the CGI fish, but also because it all happens in a lake.[9] We were supposed to start shooting now, but the longer to leave it the colder the water gets.[10] The movie takes place during Spring Break and, of course, the studio wanted it ready for the summer, but if you've got 1,000 people who need to get murdered in the water, you have to wait for the right temperature for the water, for the weather, for everything." Shooting took place in June 2009 at Bridgewater Channel in Lake Havasu, located in Lake Havasu City, Arizona. Much of one end of the channel was blocked off for boats, some flipped over and some covered in blood. The water was also dyed red for the shooting. Aja cast for his film project Wild Wild Girls, which played bikini starlets on Lake Victoria. Due to constraints with 3D camera rigs, Aja shot Piranha in 2D and converted to 3D in post production using the reali-D conversion process developed by the company, Inner-D.Unlike other 3D converted films released in 2010, Piranha's conversion was not done as an afterthought, and it represents the first post-conversion process to be well received by critics.

 

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